Secale cereale 14 Tricepiro Triticosecale 14 Univalents 8 hybrid 8 Triticum aestivum 6 article 6 chromosome 6 fluorescence in situ hybridization 6 plant chromosome 6 DNA 4 Heterochromatin 4 Hybrids 4 Meiosis 4 Micronuclei 4 Reductional division 4 Shriveled kernels 4 Thinopyrum 4 allopolyploidy 4 anaphase 4 chromatid 4 chromosome analysis 4 chromosome number 4 cultivar 4 genome 4 genomic DNA 4 genomic in situ hybridization 4 meiosis 4 nonhuman 4 rye 4