The retinal basis of light aversion in neonatal mice

Aversive responses to bright light (photoaversion) require signaling from the eye to the brain. Melanopsin-expressing intrinsically photosensitive retinal ganglion cells (ipRGCs) encode absolute light intensity and are thought to provide the light signals for photoaversion. Consistent with this, ne...

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Autores principales: Caval Holme, Franklin S., Aranda, Marcos L., Chen, Andy Q., Tiriac, Alexandre, Zhang, Yizhen, Smith, Benjamin, Birnbaumer, Lutz, Schmidt, Tiffany M., Feller, Marla B.
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Lenguaje:Inglés
Publicado: Society for Neuroscience 2023
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Acceso en línea:https://repositorio.uca.edu.ar/handle/123456789/17413
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spelling I33-R139-123456789-174132023-11-23T18:13:44Z The retinal basis of light aversion in neonatal mice Caval Holme, Franklin S. Aranda, Marcos L. Chen, Andy Q. Tiriac, Alexandre Zhang, Yizhen Smith, Benjamin Birnbaumer, Lutz Schmidt, Tiffany M. Feller, Marla B. RETINA DESARROLLO ENUCLEACION CORRIENTE FOTOELECTRICA FOTOFOBIA Aversive responses to bright light (photoaversion) require signaling from the eye to the brain. Melanopsin-expressing intrinsically photosensitive retinal ganglion cells (ipRGCs) encode absolute light intensity and are thought to provide the light signals for photoaversion. Consistent with this, neonatal mice exhibit photoaversion before the developmental onset of image vision, and melanopsin deletion abolishes photoaversion in neonates. It is not well understood how the population of ipRGCs, which constitutes multiple physiologically distinct types (denoted M1-M6 in mouse), encodes light stimuli to produce an aversive response. Here, we provide several lines of evidence that M1 ipRGCs that lack the Brn3b transcription factor drive photoaversion in neonatal mice. First, neonatal mice lacking TRPC6 and TRPC7 ion channels failed to turn away from bright light, while two photon Ca21 imaging of their acutely isolated retinas revealed reduced photosensitivity in M1 ipRGCs, but not other ipRGC types. Second, mice in which all ipRGC types except for Brn3bnegative M1 ipRGCs are ablated exhibited normal photoaversion. Third, pharmacological blockade or genetic knockout of gap junction channels expressed by ipRGCs, which reduces the light sensitivity of M2-M6 ipRGCs in the neonatal retina, had small effects on photoaversion only at the brightest light intensities. Finally, M1s were not strongly depolarized by spontaneous retinal waves, a robust source of activity in the developing retina that depolarizes all other ipRGC types. M1s therefore constitute a separate information channel between the neonatal retina and brain that could ensure behavioral responses to light but not spontaneous retinal waves... 2023-11-06T23:14:45Z 2023-11-06T23:14:45Z 2022 Artículo Caval Holme, F. S. The retinal basis of light aversion in neonatal mice [en línea]. The Journal of Neuroscience. 2022, 42(20) 4101-4115. doi: 10.1523/JNEUROSCI.0151-22.2022. Disponible en: https://repositorio.uca.edu.ar/handle/123456789/17413 0270-6474 (impreso) 1529-2401 (online) https://repositorio.uca.edu.ar/handle/123456789/17413 10.1523/JNEUROSCI.0151-22.2022 35396331 eng Acceso abierto http://creativecommons.org/licenses/by-nc-sa/4.0/ application/pdf Society for Neuroscience The Journal of Neuroscience. Vol.42, No.20, 4101-4115, 2022
institution Universidad Católica Argentina
institution_str I-33
repository_str R-139
collection Repositorio Institucional de la Universidad Católica Argentina (UCA)
language Inglés
topic RETINA
DESARROLLO
ENUCLEACION
CORRIENTE FOTOELECTRICA
FOTOFOBIA
spellingShingle RETINA
DESARROLLO
ENUCLEACION
CORRIENTE FOTOELECTRICA
FOTOFOBIA
Caval Holme, Franklin S.
Aranda, Marcos L.
Chen, Andy Q.
Tiriac, Alexandre
Zhang, Yizhen
Smith, Benjamin
Birnbaumer, Lutz
Schmidt, Tiffany M.
Feller, Marla B.
The retinal basis of light aversion in neonatal mice
topic_facet RETINA
DESARROLLO
ENUCLEACION
CORRIENTE FOTOELECTRICA
FOTOFOBIA
description Aversive responses to bright light (photoaversion) require signaling from the eye to the brain. Melanopsin-expressing intrinsically photosensitive retinal ganglion cells (ipRGCs) encode absolute light intensity and are thought to provide the light signals for photoaversion. Consistent with this, neonatal mice exhibit photoaversion before the developmental onset of image vision, and melanopsin deletion abolishes photoaversion in neonates. It is not well understood how the population of ipRGCs, which constitutes multiple physiologically distinct types (denoted M1-M6 in mouse), encodes light stimuli to produce an aversive response. Here, we provide several lines of evidence that M1 ipRGCs that lack the Brn3b transcription factor drive photoaversion in neonatal mice. First, neonatal mice lacking TRPC6 and TRPC7 ion channels failed to turn away from bright light, while two photon Ca21 imaging of their acutely isolated retinas revealed reduced photosensitivity in M1 ipRGCs, but not other ipRGC types. Second, mice in which all ipRGC types except for Brn3bnegative M1 ipRGCs are ablated exhibited normal photoaversion. Third, pharmacological blockade or genetic knockout of gap junction channels expressed by ipRGCs, which reduces the light sensitivity of M2-M6 ipRGCs in the neonatal retina, had small effects on photoaversion only at the brightest light intensities. Finally, M1s were not strongly depolarized by spontaneous retinal waves, a robust source of activity in the developing retina that depolarizes all other ipRGC types. M1s therefore constitute a separate information channel between the neonatal retina and brain that could ensure behavioral responses to light but not spontaneous retinal waves...
format Artículo
author Caval Holme, Franklin S.
Aranda, Marcos L.
Chen, Andy Q.
Tiriac, Alexandre
Zhang, Yizhen
Smith, Benjamin
Birnbaumer, Lutz
Schmidt, Tiffany M.
Feller, Marla B.
author_facet Caval Holme, Franklin S.
Aranda, Marcos L.
Chen, Andy Q.
Tiriac, Alexandre
Zhang, Yizhen
Smith, Benjamin
Birnbaumer, Lutz
Schmidt, Tiffany M.
Feller, Marla B.
author_sort Caval Holme, Franklin S.
title The retinal basis of light aversion in neonatal mice
title_short The retinal basis of light aversion in neonatal mice
title_full The retinal basis of light aversion in neonatal mice
title_fullStr The retinal basis of light aversion in neonatal mice
title_full_unstemmed The retinal basis of light aversion in neonatal mice
title_sort retinal basis of light aversion in neonatal mice
publisher Society for Neuroscience
publishDate 2023
url https://repositorio.uca.edu.ar/handle/123456789/17413
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